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The Know tool provides a structured framework for managing the Specification Graph, which serves as the single source of truth for complex software systems. By representing system components as interconnected nodes in a Directed Acyclic Graph (DAG), it ensures that all relationships—from high-level user objectives to granular implementation operations—are explicit and verifiable. This skill allows Claude to navigate the project's 'WHAT' and 'HOW' chains, perform gap analysis to identify missing dependencies, and enforce architectural rules, making it indispensable for maintaining large-scale codebases where implied relationships often lead to technical debt.