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The Performance Optimization skill provides Claude with a rigorous framework for identifying and resolving bottlenecks across the entire software stack. By emphasizing a 'measure-first' methodology, it guides developers through profiling JavaScript, Python, and database queries while implementing best practices like lazy loading, bundle size reduction, and multi-level caching strategies. It is particularly useful when scaling applications, meeting Core Web Vitals targets, or reducing infrastructure costs through efficient resource utilization.