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The Performance Optimization skill transforms Claude into a specialized performance engineer, providing a structured 'Measure First' methodology for diagnosing and resolving software slowness. It offers actionable guidance for profiling Node.js and Python environments, optimizing frontend rendering and bundle sizes, and streamlining backend database queries to eliminate N+1 problems. By integrating industry-standard metrics like Core Web Vitals and API latency targets, this skill ensures that every code change is data-driven and results in measurable improvements to the user experience.